Where do allele frequencies come from?

Mj= S(Ti*B[N-i,j-i,F]); Mj= S(Mi*B[i,i-j,R])

i=0,j i=j,N

Sj= S(Mi*B[N-i,j-i,1-1/w]); Sj= S(Mi*B[i,i-j,1-w]);

i=1,j if wɭ i=j,N-1 if wə

Dj= S Si*B[N,j,i/N]

i=1,N-1

w=relative fitness of i mutants to N-i original

Ti,Mi,Di,Si = frequency of i mutants in a pop. size N

F= forward mutation(or migration) probability ; R=reverse.

B(N,i,p)= Binomial = C(N,i) pi (1-p)N-i