If the first column contains a "-", then only known pairs of sites are involved in the clusters at this spacing. For other entries, predictions are numbered, ranked according to their probability.
For predictions based on individual separation distances, this is the most significant base-pair separation between sites.
This is the number of known pairs separated by the given spacing or spacing range.
This is the number of pairs observed at the given spacing. If this number is equal to the number of known pairs (#known), then only the known pairs were observed at the given spacing (no new predicted sites involved).
The number of pairs of sites expected by chance.
The ratio of the number of pairs observed to the number of pairs of sites expected by chance.
The probability of seeing the observed number of sites at this separation or within this separation range.